Gene Details:
- MSU gene ID: LOC_Os11g36440
- RAPdb gene ID: Os11g0572700
- Gene Symbol: OsCRTISO ZEBRA2 MHZ5 MIT3
- Genome: MSU7 , IRGSP-1.0
- Species: Oryza sativa
Functional Descriptions:
- The greatly increased activities of reactive oxygen species (ROS) scavenging enzymes, and reduced photosystem (PS) II core proteins CP43, CP47 and D1 in leaves of the OsCRTISO/phs3-1mutant and OsLCY RNAi transgenic rice indicated that photo-oxidative damage occurred in PS II, consistent with the accumulation of ROS in these plants.
- Expression analysis via both RT-PCR and ZEBRA2-promoter-beta-glucuronidase (GUS) transgenic rice indicates that ZEBRA2 is predominantly expressed in mesophyll cells of mature leaves where active photosynthesis occurs.
- More severe phenotype of the ZEBRA2 mutant under high light intensity indicates that ““zebra”” phenotype might be caused by photooxidative damages.
- We found that MHZ5 encodes a carotenoid isomerase and that the mutation in MHZ5 blocks carotenoid biosynthesis, reduces ABA accumulation, and promotes ethylene production in etiolated seedlings.
- ABA can largely rescue the ethylene response of the MHZ5 mutant.
- Ethylene induces MHZ5 expression, the production of neoxanthin, an ABA biosynthesis precursor, and ABA accumulation in roots.
- MHZ5 overexpression results in enhanced ethylene sensitivity in roots and reduced ethylene sensitivity in coleoptiles.
- Prolycopene was accumulated while all-trans-lycopene was barely detectable in the dark-grown MIT3 seedlings.
- Exogenously applied GR24, a synthetic SL, could rescue the tillering phenotype of MIT3.
- Our results reveal that the tillering phenotype of MIT3 is due to SL deficiency and directly link carotenoid deficiency with SL-regulated rice tillering.
- Mutations in the MIT3 gene encoding a caroteniod isomerase lead to increased tiller number in rice.
- Double mutant analysis of MIT3 with the SLs biosynthesis mutant d17 revealed that MIT3 controls tiller development upstream of the SLs biosynthesis pathway.
- Map-based cloning revealed that MIT3 encodes a carotenoid isomerase (CRTISO), the key enzyme catalyzing the conversion from prolycopene to all-trans-lycopene in carotenoid biosynthesis.
Function-related keywords:
- oxidative , photosynthesis , ABA , ethylene , ethylene-response , seedlings , ethylene-production , tillering , tiller , development , map-based-cloning , tiller-number
Literature:
- ZEBRA2, encoding a carotenoid isomerase, is involved in photoprotection in rice . DOI: 10.1007/s11103-010-9719-z ; PMID: 21161331
- Mutations of genes in synthesis of the carotenoid precursors of ABA lead to pre-harvest sprouting and photo-oxidation in rice . DOI: 10.1111/j.1365-313X.2008.03411.x ; PMID: 18208525
- Ethylene responses in rice roots and coleoptiles are differentially regulated by a carotenoid isomerase-mediated abscisic acid pathway . DOI: 10.1105/tpc.15.00080 ; PMID: 25841037
- Mutations in the MIT3 gene encoding a caroteniod isomerase lead to increased tiller number in rice . DOI: 10.1016/j.plantsci.2017.11.001 ; PMID: 29362087
Related News:
Gene Resources:
- NCBI ID: EF417892
- UniProt accessions:
Sequences:
cDNA Sequence
- >LOC_Os11g36440.1
GCAGATTCCACCGCGGCCTCTCCTCCCATATAAAAACCCCACCCAGCTCGAGCTCCCGCGCTTCACCGGCGCCGCCATGCCGCTCCTCCTCCTCGCCGCCGCGCGCCCGTGCGCCCCGCTCCTCGCTCCCTCCGCCGCCGGAGCAGTAGTAGGCCGGCCGTCCACCGCCCGCCCACTGGGACGAGGGACCAGGAGAGGCGCGGGCGCGGCGGCGGCGGCTGCGGTGGCGGCGGAGAAGACGGTGGTGAAGGCGGAGGAGGAGGAGGAGGGTGGGGGAGAGTACGACGCGATCGTGGTGGGGTCGGGGATCGGGGGGATGGTGGCGGCGACGCAGCTGGCGGCGAAGGGGGCGAGGGTGCTGGTGCTGGAGAAGTACGTCATCCCCGGGGGGAGCTCCGGGTACTACCGCCGCGACGGGTTCACCTTCGACGTCGGCTCCTCCGTCATGTTCGGCTTCTCCGACAAGGGGAACTTGAATTTGATTACACAAGCACTAGAAGCAGTTGGGCATAAGATGGAAGTTATACCGGATCCTTCTACGGTTCATTTCCACCTACCTGGTGACCTCTCAGTTCTTGTGCATAGAGAGTATGATGACTTCGTCACGGAGCTGGTTAATAAATTTCCTCATGAAAAGGAAGGAATCCTGAAATTCTATGGCACATGTTGGAAGATCTTCAATTCATTAAATTCTTTGGAGCTGAAGTCCCTTGAGGAGCCTCTATATCTTTTTGGGCAATTTTTTCAAAAGCCTCTGGAATGTTTGACGCTAGCTTATTATCTGCCACAGAATGCTGGGGATATTGCTCGCAAGTTCATAAAAGATCAGCAGTTACTCTCCTTCATAGATGCTGAGTGCTTTATTGTGAGCACAGTCAATGCCTTGCAAACACCCATGATTAATGCAAGCATGGTCTTGTGCGATAGGCACTTCGGGGGAATTAACTATCCAGTTGGTGGTGTTGGTGGTATTGCTATGTCCTTGGCAGATGGGCTTGTTGACAAAGGCAGTGAAATACGCTACAAGGCTAATGTGACCAATGTAATTTTAGAAAATGGCAAAGCTGTTGGAGTCAGGTTATCAAATGGGAAGGAGTTCTTTGCTAAAACAGTGATATCGAATGCTACCAGATGGGATACATTTGGTAAACTCTTGAAAGTTGAAGAGCTTCCAGAAGAGGAGAAAAACTTCCAAAAGAATTATGTTAAAGCACCATCATTCCTTTCCATTCATATGGGTGTCAAAGCCTCAGTTTTGCCTGCTGATACTGATTGCCACCATTTTGTACTTGAGGATGACTGGGCTAATTTGGAGAAGCCTTATGGAAGCATATTTTTAAGTATCCCTACAGTGCTTGACCCATCCTTGGCTCCCGAAGGACACCATATACTTCATATTTTTACAACTGCAGGGATAGAGGATTGGGAAGGTCTGTCCAGGAAGGATTATGAGAAGAAAAAGGAGCTTGTGGCGACAGAGATAATAAAGAGGCTTGAAAAAAAGCTGTTCCCTGGTCTTCAGGATTCAATAGTCCTCAAGGAGGTAGGCTCACCAAAAACACACCGAAGATTTCTTGCGCGAAATGATGGTACATATGGACCCATGCCACGGGGTAAACCAAAGGGATTGCTAGCAATGCCTTTCAATACTACTTCAATAGATGGCTTGTACTGTGTTGGCGACAGCTGTTTTCCTGGGCAAGGTGTAATTGCTGTGGCATTTTCAGGGATCATGTGTGCTCATCGAGTTGCAGCTGATATTGGACTGGAACAAAGATCTCCTGTACTAGATGCGGGTCTTCTTGGGCTCCTCAGATGGTTGAGAACGCTTGCTTAGGATGATACGGACAAAGATTTGGTTGACCGTCGATTGATTTTACGATTACAGAAAAACACCTTCTCACACATGTCATGGTCATTGCACATTTGTTTTGGTGAGCCATACATGGATCAAGCAGCAGTTTTTGCCTCTTGGGATCTTGCTTCTGCCTGGTATATACAAGAGGTTGGATTTTCAGCTGCAGTGATCCCTGAAAAGCTGAAGCTTCCTGGAATAATTGTACAGATTTCAGTTGTTATTTGGGAGCAAAAGGCGTAAAGACTAATATGGATGCATATATGTACCTAGAGCCTTATACACATGATGTAACTTGTTGTAATCTACTAGTATCTCCTGTATGCCCATGTTGTCATGTTGACAATTGAGAGTTACAAACATGCAGCTGTACATGATGTGTAAGCCTTAACAGAACAAAGTCAGATACACCTGCAGCCTTCTG
CDS Sequence
- >LOC_Os11g36440.1
ATGCCGCTCCTCCTCCTCGCCGCCGCGCGCCCGTGCGCCCCGCTCCTCGCTCCCTCCGCCGCCGGAGCAGTAGTAGGCCGGCCGTCCACCGCCCGCCCACTGGGACGAGGGACCAGGAGAGGCGCGGGCGCGGCGGCGGCGGCTGCGGTGGCGGCGGAGAAGACGGTGGTGAAGGCGGAGGAGGAGGAGGAGGGTGGGGGAGAGTACGACGCGATCGTGGTGGGGTCGGGGATCGGGGGGATGGTGGCGGCGACGCAGCTGGCGGCGAAGGGGGCGAGGGTGCTGGTGCTGGAGAAGTACGTCATCCCCGGGGGGAGCTCCGGGTACTACCGCCGCGACGGGTTCACCTTCGACGTCGGCTCCTCCGTCATGTTCGGCTTCTCCGACAAGGGGAACTTGAATTTGATTACACAAGCACTAGAAGCAGTTGGGCATAAGATGGAAGTTATACCGGATCCTTCTACGGTTCATTTCCACCTACCTGGTGACCTCTCAGTTCTTGTGCATAGAGAGTATGATGACTTCGTCACGGAGCTGGTTAATAAATTTCCTCATGAAAAGGAAGGAATCCTGAAATTCTATGGCACATGTTGGAAGATCTTCAATTCATTAAATTCTTTGGAGCTGAAGTCCCTTGAGGAGCCTCTATATCTTTTTGGGCAATTTTTTCAAAAGCCTCTGGAATGTTTGACGCTAGCTTATTATCTGCCACAGAATGCTGGGGATATTGCTCGCAAGTTCATAAAAGATCAGCAGTTACTCTCCTTCATAGATGCTGAGTGCTTTATTGTGAGCACAGTCAATGCCTTGCAAACACCCATGATTAATGCAAGCATGGTCTTGTGCGATAGGCACTTCGGGGGAATTAACTATCCAGTTGGTGGTGTTGGTGGTATTGCTATGTCCTTGGCAGATGGGCTTGTTGACAAAGGCAGTGAAATACGCTACAAGGCTAATGTGACCAATGTAATTTTAGAAAATGGCAAAGCTGTTGGAGTCAGGTTATCAAATGGGAAGGAGTTCTTTGCTAAAACAGTGATATCGAATGCTACCAGATGGGATACATTTGGTAAACTCTTGAAAGTTGAAGAGCTTCCAGAAGAGGAGAAAAACTTCCAAAAGAATTATGTTAAAGCACCATCATTCCTTTCCATTCATATGGGTGTCAAAGCCTCAGTTTTGCCTGCTGATACTGATTGCCACCATTTTGTACTTGAGGATGACTGGGCTAATTTGGAGAAGCCTTATGGAAGCATATTTTTAAGTATCCCTACAGTGCTTGACCCATCCTTGGCTCCCGAAGGACACCATATACTTCATATTTTTACAACTGCAGGGATAGAGGATTGGGAAGGTCTGTCCAGGAAGGATTATGAGAAGAAAAAGGAGCTTGTGGCGACAGAGATAATAAAGAGGCTTGAAAAAAAGCTGTTCCCTGGTCTTCAGGATTCAATAGTCCTCAAGGAGGTAGGCTCACCAAAAACACACCGAAGATTTCTTGCGCGAAATGATGGTACATATGGACCCATGCCACGGGGTAAACCAAAGGGATTGCTAGCAATGCCTTTCAATACTACTTCAATAGATGGCTTGTACTGTGTTGGCGACAGCTGTTTTCCTGGGCAAGGTGTAATTGCTGTGGCATTTTCAGGGATCATGTGTGCTCATCGAGTTGCAGCTGATATTGGACTGGAACAAAGATCTCCTGTACTAGATGCGGGTCTTCTTGGGCTCCTCAGATGGTTGAGAACGCTTGCTTAG
Protein Sequence
- >LOC_Os11g36440.1
MPLLLLAAARPCAPLLAPSAAGAVVGRPSTARPLGRGTRRGAGAAAAAAVAAEKTVVKAEEEEEGGGEYDAIVVGSGIGGMVAATQLAAKGARVLVLEKYVIPGGSSGYYRRDGFTFDVGSSVMFGFSDKGNLNLITQALEAVGHKMEVIPDPSTVHFHLPGDLSVLVHREYDDFVTELVNKFPHEKEGILKFYGTCWKIFNSLNSLELKSLEEPLYLFGQFFQKPLECLTLAYYLPQNAGDIARKFIKDQQLLSFIDAECFIVSTVNALQTPMINASMVLCDRHFGGINYPVGGVGGIAMSLADGLVDKGSEIRYKANVTNVILENGKAVGVRLSNGKEFFAKTVISNATRWDTFGKLLKVEELPEEEKNFQKNYVKAPSFLSIHMGVKASVLPADTDCHHFVLEDDWANLEKPYGSIFLSIPTVLDPSLAPEGHHILHIFTTAGIEDWEGLSRKDYEKKKELVATEIIKRLEKKLFPGLQDSIVLKEVGSPKTHRRFLARNDGTYGPMPRGKPKGLLAMPFNTTSIDGLYCVGDSCFPGQGVIAVAFSGIMCAHRVAADIGLEQRSPVLDAGLLGLLRWLRTLA*