Gene Details:
- MSU gene ID: LOC_Os05g44760
- RAPdb gene ID: Os05g0522500
- Gene Symbol: OsHXK5
- Genome: MSU7 , IRGSP-1.0
- Species: Oryza sativa
Functional Descriptions:
- Notably, the expression of OsHXK5, OsHXK6, or their mutant alleles complemented the Arabidopsis glucose insensitive2-1 mutant, thereby resulting in wild-type characteristics in seedling development, Glc-dependent gene expression, and plant growth.
- Furthermore, transgenic rice plants overexpressing OsHXK5 or OsHXK6 exhibited hypersensitive plant growth retardation and enhanced repression of the photosynthetic gene RbcS in response to Glc treatment.
- Transient expression analyses using GFP fusion constructs revealed that OsHXK5 and OsHXK6 are associated with mitochondria.
- Interestingly, the OsHXK5DeltamTP-GFP and OsHXK6DeltamTP-GFP fusion proteins, which lack N-terminal mitochondrial targeting peptides, were present mainly in the nucleus with a small amount of the proteins seen in the cytosol.
- In addition, the OsHXK5NLS-GFP and OsHXK6NLS-GFP fusion proteins harboring nuclear localization signals were targeted predominantly in the nucleus, suggesting that these OsHXKs retain a dual-targeting ability to mitochondria and nuclei.
- Sugars induced the expression of three OsHXKs, OsHXK2, OsHXK5, and OsHXK6, in excised leaves, while suppressing OsHXK7 expression in excised leaves and immature seeds.
- Our results demonstrate that OsHXK5 contributes to a large portion of the hexokinase activity necessary for the starch utilization pathway during pollen germination and tube growth, as well as for starch biosynthesis during pollen maturation.
- We characterized hxk5-1, a Tos17 mutant of OsHXK5 that is up-regulated in maturing pollen, a stage when starch accumulates.
Function-related keywords:
- seedling , growth , mitochondria , seed , pollen , starch , starch-biosynthesis
Literature:
- Structure, expression, and functional analysis of the hexokinase gene family in rice (Oryza sativa L.) . DOI: 10.1007/s00425-006-0251-y ; PMID: 16552590
- Role of the rice hexokinases OsHXK5 and OsHXK6 as glucose sensors . DOI: 10.1104/pp.108.131227 ; PMID: 19010999
- Deficiency of rice hexokinase HXK5 impairs synthesis and utilization of starch in pollen grains and causes male sterility . DOI: 10.1093/jxb/erz436 ; PMID: 31671177
Related News:
Gene Resources:
- NCBI ID: DQ116387
- UniProt accessions:
Sequences:
cDNA Sequence
- >LOC_Os05g44760.1
ACCGGTTAAAACGCCCAAGTCAGTGGCTCTCGTCCTCCTTTCTCCTACGCGGCCGGAGAGAGGAGGGGGGAAGAGAGGATCTCGCAGAGCGCCATAGCCTCGGATCCAGAAGCGGAATTCGGTGGAGGTCTCGGGCACCTGGATCGATCGGAGGAAGGGAAGGCGGAGCAGCGGTGATGGGGAAGGCGGCGGCGGTGGGGACGGCGGTGGTGGTGGCCGCGGCGGTCGGGGTGGCGGTGGTGCTGGCGCGGAGGCGGAGGAGGAGGGACCTGGAGCTGGTGGAGGGAGCCGCGGCGGAGAGGAAGAGGAAGGTGGCGGCGGTGATCGAGGACGTGGAGCACGCGCTGTCGACCCCGACGGCGCTGCTGCGGGGCATCTCGGACGCCATGGTCACCGAGATGGAGCGAGGCCTGCGCGGGGACAGCCACGCCATGGTTAAGATGCTCATCACCTACGTCGACAACCTCCCCACCGGAAATGAACAGGGGTTGTTTTATGCATTGGATCTTGGAGGAACCAACTTCCGCGTCCTGCGAGTCCAACTCGGTGGCAAGGAGAAACGTGTCGTCCAACAACAGTACGAGGAAGTCTCAATTCCACCACATCTGATGGTTGGAACTTCCATGGAACTGTTTGATTTTATTGCTTCTGCATTGTCAAAATTTGTTGATACTGAAGGTGATGATTTCCACCTCCCAGAAGGGAGACAGAGAGAGCTGGGCTTCACCTTTTCCTTCCCAGTGAGCCAGACATCAATATCGTCAGGAACGCTCATCAAGTGGACAAAGGGTTTCTCCATCAATGACGCGGTTGGCGAAGATGTTGTATCTGAGTTGGGCAAGGCCATGGAGAGGCAGGGATTAGATATGAAAATTGCAGCATTGGTTAATGACACTGTCGGCACATTGGCTGGTGGGAGGTATGCGGATAACAGTGTTGTTGCTGCTATAATATTGGGCACTGGTACAAATGCAGCATATGTTGAGAATGCTAATGCAATTCCTAAATGGACCGGTTTACTGCCTAGGTCCGGAAATATGGTAATCAACACAGAATGGGGGAGCTTTAAATCAGATAAGCTTCCTCTTTCAGAATTCGATAAAGCAATGGATTTTGAAAGCTTGAATCCTGGAGAGCAGATATATGAAAAGTTGATTTCTGGCATGTATCTTGGAGAGATAGTGCGAAGAATCTTGCTGAAGCTTGCTCATGATGCAGCTTTGTTTGGGGATGTTGTTCCATCTAAGCTAGAGCAACCGTTTGTACTAAGGACACCGGATATGTCAGCCATGCATCATGACTCGTCACATGACCTTAAAACTGTTGGAGCTAAGCTAAAGGATATCGTCGGGGTCCCAGATACTTCCCTGGAAGTAAGATACATTACCAGTCACATTTGTGACATAGTTGCAGAGCGTGCTGCACGCTTGGCTGCTGCTGGCATATATGGGGTCCTAAAGAAGCTAGGTCGGGACAAGATGCCAAAAGACGGCAGTAAGATGCCTAGGACTGTCATTGCCTTGGATGGTGGGCTCTATGAACATTACAAGAAGTTCAGCAGTTGCTTAGAATCAACTCTAACAGACCTTCTTGGGGATGATGTCTCGTCTTCGGTGGTTACCAAGCTGGCCAACGATGGTTCTGGCATTGGAGCTGCTCTTCTCGCAGCCTCGCACTCCCAGTATGCCGAGATCGACTAGCTTTAAGGATGATCTTGATGAATGATGAATCAAACTCCGTTTGTAGGTTCTCATTTCCCCCTTCAAAATCCACATAATACTCCTGGCTCCCCCCTTGAAATCTTACCATCTTTTTTTGGCTATTCTGAGGGCAAACATAAGTGCCTCTGCAGCGGGATATAGCTAGTATAGCGCCAATGAGTTTGGAGGTTTTCTAATGGCATAAAACGTTGGATGGCAGTAGCAGACTAACAGGGAAATGGAGGCACAGGCAATTTCCATTCCTGTTCTGTCAGATTCTTTTCCCCCTTAATTGATGTTGAGAACCAAGATTTTTTTGCTCTGTATTTTCTCTTCGTAATAAAGAAGGGGACATAATCTAATTGCTCTTGTTTGATCTCATATATGTATGTGACTGCACTCTCACTGTCCCAAAATTTTATAGATGCATTTACAAAGTTACAATACTCATTACATGCAAATAACTGAAGGATGGCGAATGTATTTGTCTGGTATCTACCTGCTCTCCTAACTAGCGTGCCATCTCTCCTAGATGGTAACCATTGGATCAGGTTATACTTCTTATTGACCATTGTTCTCATACTATGCTGTCCAAGCTGCCGTTGCCGTCGTCGATGCCTTTGGTGTCCGAAAGCTGCTTCCGGCGGCTGCCACCCCTGATCGGCAGAAACGGCGCGCCGTCGCCGTCGTACTCCGCCACCTTCCCGTCCACGGTGTTCTGGAGCGCCTGCTCGCACCTCACCAGAATCTGCAGAACGTCCCTCATTGTCGGCCTCGTCGCCGGCTGCGCGCCGGTGCAGATGATGCCGAGCTTGAACACGACCTCGGCGTCGTCGCCGTACCCGGAGTCGGTGATGCACCTGTCCACCGCGTCGGCGATGCTCCTGCCCGATTGCAGGTGCCGCCACGCCCACTCCGCCAGCGAGCCGTGCTCGCCGCCGTCGTGGGCTTCCCTGCCGGTGATCAGCTCGAGGAGCACCACCCCGAAGCTGTACACGTCGACCTTCTCGTTCACCTTCCTCGTGTACGCGCACTCTGCA
CDS Sequence
- >LOC_Os05g44760.1
ATGGGGAAGGCGGCGGCGGTGGGGACGGCGGTGGTGGTGGCCGCGGCGGTCGGGGTGGCGGTGGTGCTGGCGCGGAGGCGGAGGAGGAGGGACCTGGAGCTGGTGGAGGGAGCCGCGGCGGAGAGGAAGAGGAAGGTGGCGGCGGTGATCGAGGACGTGGAGCACGCGCTGTCGACCCCGACGGCGCTGCTGCGGGGCATCTCGGACGCCATGGTCACCGAGATGGAGCGAGGCCTGCGCGGGGACAGCCACGCCATGGTTAAGATGCTCATCACCTACGTCGACAACCTCCCCACCGGAAATGAACAGGGGTTGTTTTATGCATTGGATCTTGGAGGAACCAACTTCCGCGTCCTGCGAGTCCAACTCGGTGGCAAGGAGAAACGTGTCGTCCAACAACAGTACGAGGAAGTCTCAATTCCACCACATCTGATGGTTGGAACTTCCATGGAACTGTTTGATTTTATTGCTTCTGCATTGTCAAAATTTGTTGATACTGAAGGTGATGATTTCCACCTCCCAGAAGGGAGACAGAGAGAGCTGGGCTTCACCTTTTCCTTCCCAGTGAGCCAGACATCAATATCGTCAGGAACGCTCATCAAGTGGACAAAGGGTTTCTCCATCAATGACGCGGTTGGCGAAGATGTTGTATCTGAGTTGGGCAAGGCCATGGAGAGGCAGGGATTAGATATGAAAATTGCAGCATTGGTTAATGACACTGTCGGCACATTGGCTGGTGGGAGGTATGCGGATAACAGTGTTGTTGCTGCTATAATATTGGGCACTGGTACAAATGCAGCATATGTTGAGAATGCTAATGCAATTCCTAAATGGACCGGTTTACTGCCTAGGTCCGGAAATATGGTAATCAACACAGAATGGGGGAGCTTTAAATCAGATAAGCTTCCTCTTTCAGAATTCGATAAAGCAATGGATTTTGAAAGCTTGAATCCTGGAGAGCAGATATATGAAAAGTTGATTTCTGGCATGTATCTTGGAGAGATAGTGCGAAGAATCTTGCTGAAGCTTGCTCATGATGCAGCTTTGTTTGGGGATGTTGTTCCATCTAAGCTAGAGCAACCGTTTGTACTAAGGACACCGGATATGTCAGCCATGCATCATGACTCGTCACATGACCTTAAAACTGTTGGAGCTAAGCTAAAGGATATCGTCGGGGTCCCAGATACTTCCCTGGAAGTAAGATACATTACCAGTCACATTTGTGACATAGTTGCAGAGCGTGCTGCACGCTTGGCTGCTGCTGGCATATATGGGGTCCTAAAGAAGCTAGGTCGGGACAAGATGCCAAAAGACGGCAGTAAGATGCCTAGGACTGTCATTGCCTTGGATGGTGGGCTCTATGAACATTACAAGAAGTTCAGCAGTTGCTTAGAATCAACTCTAACAGACCTTCTTGGGGATGATGTCTCGTCTTCGGTGGTTACCAAGCTGGCCAACGATGGTTCTGGCATTGGAGCTGCTCTTCTCGCAGCCTCGCACTCCCAGTATGCCGAGATCGACTAG
Protein Sequence
- >LOC_Os05g44760.1
MGKAAAVGTAVVVAAAVGVAVVLARRRRRRDLELVEGAAAERKRKVAAVIEDVEHALSTPTALLRGISDAMVTEMERGLRGDSHAMVKMLITYVDNLPTGNEQGLFYALDLGGTNFRVLRVQLGGKEKRVVQQQYEEVSIPPHLMVGTSMELFDFIASALSKFVDTEGDDFHLPEGRQRELGFTFSFPVSQTSISSGTLIKWTKGFSINDAVGEDVVSELGKAMERQGLDMKIAALVNDTVGTLAGGRYADNSVVAAIILGTGTNAAYVENANAIPKWTGLLPRSGNMVINTEWGSFKSDKLPLSEFDKAMDFESLNPGEQIYEKLISGMYLGEIVRRILLKLAHDAALFGDVVPSKLEQPFVLRTPDMSAMHHDSSHDLKTVGAKLKDIVGVPDTSLEVRYITSHICDIVAERAARLAAAGIYGVLKKLGRDKMPKDGSKMPRTVIALDGGLYEHYKKFSSCLESTLTDLLGDDVSSSVVTKLANDGSGIGAALLAASHSQYAEID*