Gene Details:
- MSU gene ID: LOC_Os03g46470
- RAPdb gene ID: Os03g0667500
- Gene Symbol: OsIRT1
- Genome: MSU7 , IRGSP-1.0
- Species: Oryza sativa
Functional Descriptions:
- RNA interference of OsIRO2 in transgenic rice showed that ethylene acted via this transcription factor to induce the expression of OsNAS1, OsNAS2, OsYSL15, and OsIRT1.
- This suggests that OsIRT1 is a functional metal transporter for iron, and is actively engaged in Fe uptake from soils, especially under limiting conditions.
- The knockout of OsARF12 also altered the abundance of mitochondrial iron-regulated (OsMIR), iron (Fe)-regulated transporter1 (OsIRT1) and short postembryonic root1 (OsSPR1) in roots of rice, and resulted in lower Fe content.
- In spite of being a Strategy II plant, however, rice (Oryza sativa) contains the previously identified Fe2+ transporter OsIRT1.
- Real-time PCR analysis revealed that OsIRT1 and OsIRT2 are expressed predominantly in roots, and these transporters are induced by low-Fe conditions.
- Over-expression of OsIRT1 leads to increased iron and zinc accumulations in rice.
- Northern blot analysis revealed that OsIRT1 mRNA was predominantly expressed in roots and was induced by Fe and Cu‐deficiency.
- Yeast mutants expressing OsIRT1 and OsIRT2, which encode the rice Fe2+ transporter, became more sensitive to Cd, suggesting that Cd was absorbed by OsIRT1 and OsIRT2.
- Iron deficiency enhances cadmium uptake and translocation mediated by the Fe2+transporters OsIRT1 and OsIRT2 in rice.
- A yeast Fe3+ chelate-reductase gene refre1/372, selected for better performance at high pH, was fused to the promoter of the Fe-regulated transporter, OsIRT1, and introduced into rice plants.
- We generated transgenic rice plants over-expressing OsIRT1 to evaluate its functional roles in metal homeostasis.
- OsIRT1 promoter-GUS analysis revealed that OsIRT1 is expressed in the epidermis and exodermis of the elongating zone and in the inner layer of the cortex of the mature zone of Fe-deficient roots.
- Four distinct genes, OsZIP4, OsZIP5, OsZIP6, and OsZIP7 that exhibit sequence similarity to the rice ferrous ion transporter, OsIRT1, were isolated.
- This demonstrates that OsIRT1 can be used for enhancing micronutrient levels in rice grains.
- When expressed in yeast cells, OsIRT1 cDNA reversed the growth defects of the yeast iron‐uptake mutant.
Function-related keywords:
- ethylene , transporter , root , iron , transcription-factor , homeostasis , cadmium , zinc , mitochondria , grain , growth
Literature:
- Ethylene is involved in the regulation of iron homeostasis by regulating the expression of iron-acquisition-related genes in Oryza sativa . DOI: 10.1093/jxb/erq301 ; PMID: 21112958
- Rice plants take up iron as an Fe3+-phytosiderophore and as Fe2+ . DOI: 10.1111/j.1365-313X.2005.02624.x ; PMID: 16412081
- OsZIP4, a novel zinc-regulated zinc transporter in rice . DOI: 10.1093/jxb/eri317 ; PMID: 16263903
- Cloning an iron-regulated metal transporter from rice . DOI: 10.1093/jxb/erf004 ; PMID: 12096107
- Isolation and characterization of IRO2, a novel iron-regulated bHLH transcription factor in graminaceous plants . DOI: 10.1093/jxb/erl054 ; PMID: 16887895
- A receptor-like protein RMC is involved in regulation of iron acquisition in rice . DOI: 10.1093/jxb/ert290 ; PMID: 24014863
- Over-expression of OsIRT1 leads to increased iron and zinc accumulations in rice . DOI: 10.1111/j.1365-3040.2009.01935.x ; PMID: 19183299
- The rice transcription factor IDEF1 is essential for the early response to iron deficiency, and induces vegetative expression of late embryogenesis abundant genes . DOI: 10.1111/j.1365-313X.2009.04015.x ; PMID: 19737364
- OsARF12, a transcription activator on auxin response gene, regulates root elongation and affects iron accumulation in rice (Oryza sativa) . DOI: 10.1111/j.1469-8137.2011.03910.x ; PMID: 21973088
- Iron deficiency enhances cadmium uptake and translocation mediated by the Fe2+ transporters OsIRT1 and OsIRT2 in rice . DOI: 10.1111/j.1747-0765.2006.00055.x
- Mutational reconstructed ferric chelate reductase confers enhanced tolerance in rice to iron deficiency in calcareous soil . DOI: 10.1073/pnas.0610555104 ; PMID: 17449639
- Comparative transcriptome profile analysis of rice varieties with different tolerance to zinc deficiency . DOI: 10.1111/plb.13227 ; PMID: 33296551
- Effects of Fe and Mn cations on Cd uptake by rice plant in hydroponic culture experiment . DOI: 10.1371/journal.pone.0243174 ; PMID: 33301482
Related News:
Gene Resources:
- NCBI ID: AB070226
- UniProt accessions:
Sequences:
cDNA Sequence
- >LOC_Os03g46470.1
AGCTCAAGCAAGCAGCGCGCCTGAGTACGTAGCAAAATCCAGTGTGGTGTACGAACGAAAATGGCGACGCCGCGGACACTGGTGCCCATTCTGCCGCCCGTCGCGGCGCTCCTCCTCCTCCTCGTCGCCGCCTCCTCCATCCCCATCCTCGCCGCCGCGCAGCCGGCGGACGCGTGCGGCGGCGCACCGGATCAGGCGGCGGCGGACGGCGCGTGCCACGACGTGCCGAGGGCGCTGCGGCTGAAGCTGATCGCCATCCCGACCATCCTCGTGTCGAGCGTCGTCGGCGTGTGCCTGCCGCTCCTCTCCCGCTCCGTGCCGGCGCTCCGCCCCGACGGCGGCCTCTTCGCCGTCGTCAAGGCGTTCGCGTCGGGCGTCATCCTCGCCACGGGCTACATGCACGTGCTCCCGGACGCCTTCAACAACCTCACCTCGCCGTGCCTGCCCAGGAAGCCGTGGTCGGAGTTCCCGTTCGCGGCGTTCGTCGCCATGCTCGCCGCCGTGTCCACGCTCATGGCCGACTCGCTCATGCTCACCTACTACAACCGCAGCAAGCCCCGGCCGTCTAGCGGCGGCGACGTCGCCGCCGTCGCCGACCACGGCGAGAGCCCCGACCAGGGCCACCGGCACGGACACGGACACGGCCATGGGCATGGCATGGCGGTGGCCAAGCCCGACGACGTCGAGGCCACTCAGGTGCAGCTGCGCCGGAACCGCGTCGTCGTTCAGGTCCTCGAGATAGGCATCGTGGTGCACTCGGTGGTGATCGGCCTCGGCATGGGGGCGTCGCAGAACGTGTGCACCATCCGGCCGCTGGTGGCGGCGATGTGCTTCCACCAGATGTTCGAGGGGATGGGCCTCGGCGGCTGCATCCTGCAGGCGGAGTACGGCCGCCGGATGAGGTCGGTGCTCGTCTTCTTCTTCTCCACCACGACGCCGTTCGGCATCGCCCTCGGCCTCGCCCTGACCAGGGTGTACAGGGACAACAGCCCGACGGCGCTCATCGTCGTCGGCCTCCTCAACGCCGCCTCCGCGGGGCTGCTCCACTACATGGCGCTGGTGGAGCTCCTCGCCGCCGACTTCATGGGGCCCAAGCTGCAGGGCAACGTCCGCCTCCAGCTCGCCGCCTTCCTCGCCGTCCTCCTCGGCGCCGGCGGCATGTCCGTCATGGCCAAGTGGGCGTGAGCTGACGATGCATCACACTCTGCGGCATCATGCAATTCGCTGCATTGTTAGATTGAGCTTGGTCAGACTCGATCATCAGCTGCAGACATGTTGTACAGTGACTGTGACTTCTCCATGTTGTACTGTAATTCAGGTTTTCAGTGTTGTAGCGTGGTTGTTCAGGTTTTGTTCTTCGTGACTGATGATGTACTTCAATACGAGCATCTACATATTTTCATGATTATTACATTATCAAGCCAAAGAACTATGGTAAGATGGCAAATTTCACACAATATGACAATGTCAACAGGAATGGATGAAAATATGGAATCATGAAAAATTTC
CDS Sequence
- >LOC_Os03g46470.1
ATGGCGACGCCGCGGACACTGGTGCCCATTCTGCCGCCCGTCGCGGCGCTCCTCCTCCTCCTCGTCGCCGCCTCCTCCATCCCCATCCTCGCCGCCGCGCAGCCGGCGGACGCGTGCGGCGGCGCACCGGATCAGGCGGCGGCGGACGGCGCGTGCCACGACGTGCCGAGGGCGCTGCGGCTGAAGCTGATCGCCATCCCGACCATCCTCGTGTCGAGCGTCGTCGGCGTGTGCCTGCCGCTCCTCTCCCGCTCCGTGCCGGCGCTCCGCCCCGACGGCGGCCTCTTCGCCGTCGTCAAGGCGTTCGCGTCGGGCGTCATCCTCGCCACGGGCTACATGCACGTGCTCCCGGACGCCTTCAACAACCTCACCTCGCCGTGCCTGCCCAGGAAGCCGTGGTCGGAGTTCCCGTTCGCGGCGTTCGTCGCCATGCTCGCCGCCGTGTCCACGCTCATGGCCGACTCGCTCATGCTCACCTACTACAACCGCAGCAAGCCCCGGCCGTCTAGCGGCGGCGACGTCGCCGCCGTCGCCGACCACGGCGAGAGCCCCGACCAGGGCCACCGGCACGGACACGGACACGGCCATGGGCATGGCATGGCGGTGGCCAAGCCCGACGACGTCGAGGCCACTCAGGTGCAGCTGCGCCGGAACCGCGTCGTCGTTCAGGTCCTCGAGATAGGCATCGTGGTGCACTCGGTGGTGATCGGCCTCGGCATGGGGGCGTCGCAGAACGTGTGCACCATCCGGCCGCTGGTGGCGGCGATGTGCTTCCACCAGATGTTCGAGGGGATGGGCCTCGGCGGCTGCATCCTGCAGGCGGAGTACGGCCGCCGGATGAGGTCGGTGCTCGTCTTCTTCTTCTCCACCACGACGCCGTTCGGCATCGCCCTCGGCCTCGCCCTGACCAGGGTGTACAGGGACAACAGCCCGACGGCGCTCATCGTCGTCGGCCTCCTCAACGCCGCCTCCGCGGGGCTGCTCCACTACATGGCGCTGGTGGAGCTCCTCGCCGCCGACTTCATGGGGCCCAAGCTGCAGGGCAACGTCCGCCTCCAGCTCGCCGCCTTCCTCGCCGTCCTCCTCGGCGCCGGCGGCATGTCCGTCATGGCCAAGTGGGCGTGA
Protein Sequence
- >LOC_Os03g46470.1
MATPRTLVPILPPVAALLLLLVAASSIPILAAAQPADACGGAPDQAAADGACHDVPRALRLKLIAIPTILVSSVVGVCLPLLSRSVPALRPDGGLFAVVKAFASGVILATGYMHVLPDAFNNLTSPCLPRKPWSEFPFAAFVAMLAAVSTLMADSLMLTYYNRSKPRPSSGGDVAAVADHGESPDQGHRHGHGHGHGHGMAVAKPDDVEATQVQLRRNRVVVQVLEIGIVVHSVVIGLGMGASQNVCTIRPLVAAMCFHQMFEGMGLGGCILQAEYGRRMRSVLVFFFSTTTPFGIALGLALTRVYRDNSPTALIVVGLLNAASAGLLHYMALVELLAADFMGPKLQGNVRLQLAAFLAVLLGAGGMSVMAKWA*